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Cohen, B.L. et al. (1997) Enigmatic phylogeny of skuas (Aves: Stercorariidae). Proceedings of the Royal Society of London Series B: Biological Sciences, 264(1379), pp. 181-190. (doi: 10.1098/rspb.1997.0026) (PMID:9061968) (PMCID:PMC1688246)

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